US2011143418A1PendingUtilityA1

Compositions and methods relating to orthogonal mrna pairs

Assignee: MEDICAL RES COUNCILPriority: Jul 15, 2005Filed: Dec 8, 2010Published: Jun 16, 2011
Est. expiryJul 15, 2025(expired)· nominal 20-yr term from priority
C12N 15/67C12N 15/1034C12N 15/1089C12N 15/11
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Claims

Abstract

Orthogonal ribosome orthogonal mRNA pairs are provided, as are methods for their selection involving a novel positive-negative selection approach, and methods for their use. Also provided are cellular logic circuits involving orthogonal ribosomes.

Claims

exact text as granted — not AI-modified
1 . An orthogonal mRNA orthogonal rRNA pair, wherein expression of said orthogonal rRNA is not toxic to the cell in which it is expressed. 
     
     
         2 . The orthogonal mRNA orthogonal rRNA pair of  claim 1 , wherein said orthogonal mRNA comprises a sequence in the region from −13 to −7, relative to the AUG initiation codon, selected from the group consisting of: 5′, −13 CACCACX −7 -3′ (SEQ ID NO: 48, where X is a deletion of the nucleotide at −7 relative to the AUG); 5′- −13 CAACUGC −7 -3′ (SEQ ID NO: 49); 5′-  −13 .CAUCCCU −7 -3′ (SEQ ID NO: 50); AND 5′-  −13 UCCCUXX −7 -3′ ((SEQ ID NO: 51, where X is a deletion of the nucleotides at −7 and −8 relative to the AUG). 
     
     
         3 . The orthogonal mRNA orthogonal rRNA pair of  claim 1 , wherein said orthogonal mRNA comprises a sequence immediately 5′ of the AUG initiation codon, selected from the group consisting of: 5′-CACCACCCGCAA-3′ (SEQ ID NO: 52); 5′-CAACUGCCCGCAA-3′ (SEQ ID NO: 53); 5′-CAUCCCUCCGCAA-3′ (SEQ ID NO: 54); and 5′-UCCCUCCGCAA-3′ (SEQ ID NO: 55). 
     
     
         4 . The orthogonal mRNA orthogonal rRNA pair of  claim 1 , wherein said orthogonal rRNA comprises a sequence in the regions corresponding to nucleotides 722-723 and 1535-1540 of  E. coli  16S rRNA selected from the group consisting of (a)-(j) below: 
       
         
           
                 
                 
                 
               
                     
                 
                   Orthogonal  
                     
                     
                 
                   rRNA sequence 
                   722-723 
                   1535-1540 
                 
                     
                 
                   a 
                   AG 
                   AGUGGU (SEQ ID NO: 56) 
                 
                     
                 
                   b 
                   CG 
                   CUGUGG (SEQ ID Na 57) 
                 
                     
                 
                   c 
                   CA 
                   UUGUGG (SEQ ID NO: 58) 
                 
                     
                 
                   d 
                   GU 
                   UUGUGG (SEQ ID NO: 59) 
                 
                     
                 
                   e 
                   AC 
                   AUGCAG (SEQ ID NO: 60) 
                 
                     
                 
                   f 
                   AC 
                   UUGCAG (SEQ ID NO: 61) 
                 
                     
                 
                   g 
                   CG 
                   UCGCAG (SEQ ID NO: 62) 
                 
                     
                 
                   h 
                   CA 
                   CCGCAG (SEQ ID NO: 63) 
                 
                     
                 
                   i 
                   CA 
                   UGGGAU (SEQ ID NO: 64) 
                 
                     
                 
                   j 
                   GU 
                   UGGGAU (SEQ ID NO: 65) 
                 
                     
                 
             
                
                
                
                
               
               
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         5 . The orthogonal mRNA orthogonal rRNA pair of  claim 1 , wherein said orthogonal rRNA comprises a sequence in the regions corresponding to nucleotides 719-726 and 1531-1542 of  E. coli  16S rRNA selected from the group consisting of (a)-(j) below: 
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                   Orthogonal  
                     
                     
                 
                     
                   rRNA sequence 
                   719-726 
                   1531-1542 
                 
                     
                     
                 
                     
                   a 
                   CCGAGUGGC 
                   AUCAAGUGGUUA 
                 
                     
                     
                   (SEQ ID NO: 66) 
                   (SEQ ID NO: 67) 
                 
                     
                     
                 
                     
                   b 
                   CCGCGUGGC 
                   AUCACUGUGGUA 
                 
                     
                     
                   (SEQ ID NO: 68) 
                   (SEQ ID NO: 69) 
                 
                     
                     
                 
                     
                   c 
                   CCGCAUGGC 
                   AUCAUUGUGGUA 
                 
                     
                     
                   (SEQ ID NO: 70) 
                   (SEQ ID NO: 71) 
                 
                     
                     
                 
                     
                   d 
                   CCGGUUGGC 
                   AUCAUUGUGGUA 
                 
                     
                     
                   (SEQ ID NO: 72) 
                   (SEQ ID NO: 73) 
                 
                     
                     
                 
                     
                   e 
                   CCGACUGGC 
                   AUCAAUGCAGUA 
                 
                     
                     
                   (SEQ ID NO: 74) 
                   (SEQ ID NO: 75) 
                 
                     
                     
                 
                     
                   f 
                   CCGACUGGC 
                   AUCAAUGCAGUA 
                 
                     
                     
                   (SEQ ID NO: 76) 
                   (SEQ ID NO: 77) 
                 
                     
                     
                 
                     
                   g 
                   CCGCGUGGC 
                   AUCAUCGCAGUA 
                 
                     
                     
                   (SEQ ID NO: 78) 
                   (SEQ ID NO: 79) 
                 
                     
                     
                 
                     
                   h 
                   CCGCAUGGC 
                   AUCACCGCAGUA 
                 
                     
                     
                   (SEQ ID NO: 80) 
                   (SEQ ID NO: 81) 
                 
                     
                     
                 
                     
                   i 
                   CCGCAUGGC 
                   AUCAUGGGALTUA 
                 
                     
                     
                   (SEQ ID NO: 80) 
                   (SEQ ID NO: 82) 
                 
                     
                     
                 
                     
                   j 
                   CCGGUUGGC 
                   AUCAUGGGAUUA 
                 
                     
                     
                   (SEQ ID NO: 83) 
                   (SEQ ID NO: 82) 
                 
                     
                     
                 
             
                
                
                
                
               
               
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         6 . The orthogonal mRNA orthogonal rRNA pair of  claim 1 , wherein said pair comprises sequences selected from the paired mRNA and rRNA sequences: 
       
         
           
                 
                 
                 
               
                     
                 
                     
                   Orthogonal mRNA: 
                     
                 
                     
                   −13 to −7 
                     
                 
                     
                   relative to AUG 
                   orthogonal rRNA: 
                 
                   Pair 
                   initiation codon 
                   722-723/1535-1540 
                 
                     
                 
                     
                 
                 
                 
                 
               
                   1 
                   5′- −13 CACCACX −7 -3′ 
                   5′-A 722 G 72·3/1535 AGUGGU 1540 -3′ 
                 
                     
                   (SEQ 113 NO: 84) 
                   (SEQ ID NO: 85) 
                 
                     
                 
                   2 
                   5′- −13 CACCACX −7 -3′ 
                   5′ - C 722 G 723/1535 CUGUGG 1540 -3′ 
                 
                     
                   (SEQ ID NO: 84) 
                   (SEQ ID NO: 86) 
                 
                     
                 
                   3 
                   5′- −13 CACCACX −7 -3′ 
                   5′-C 722 A 723/1535 UUGUGG 1540 -3′ 
                 
                     
                   (SEQ ID NO: 84) 
                   (SEQ ID NO: 87) 
                 
                     
                 
                   4 
                   5′- −13 CACCACX −7 -3′ 
                   5′-G 722 U 723/1535 UUGUGG 1540 -3′ 
                 
                     
                   (SEQ ID NO: 84) 
                   (SEQ ID NO: 87) 
                 
                     
                 
                   5 
                   5′- −13 CAACUGC −7 -3′ 
                   5′-A 722 U 723/1535 AUGCAG 1540 -3′ 
                 
                     
                   (SEQ ID NO: 88) 
                   (SEQ ID NO: 89) 
                 
                     
                 
                   6 
                   5′- −13 CAACUGC −7 -3′ 
                   5′-A 722 C 723/1535 UUGCAG 1540 -3′ 
                 
                     
                   (SEQ ID NO: 88) 
                   (SEQ ID NO: 90) 
                 
                     
                 
                   7 
                   5′- −13 CAACUGC −7 -3′ 
                   5 ′- C 722 G7 23/1535 UCGCAG 1540 -3′ 
                 
                     
                   (SEQ ID NO: 88) 
                   (SEQ ID NO: 91) 
                 
                     
                 
                   8 
                   5′- −13 CAACUGC −7 -3′ 
                   5 ′- C 722 A 723/1535 CCGCAG 1540 -3′ 
                 
                     
                   (SEQ ID NO: 88) 
                   (SEQ ID NO: 92) 
                 
                     
                 
                   9 
                   5′- −13 CAUCCCU −7 -3′ 
                   5 ′- C 722 A 723/1535 UGGGAU 1540 -3′ 
                 
                     
                   (SEQ ID NO: 93) 
                   (SEQ ID NO: 94) 
                 
                     
                 
                   10 
                   5′- −13 CAUCCCU -7 -3′ 
                   5 ′- C 722 U 723/1535 UGGGAU 1540 -3′ 
                 
                     
                   (SEQ ID NO: 93) 
                   (SEQ ID NO: 94) 
                 
                     
                 
             
                
                
                
                
                
                
               
               
                
               
            
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         7 . The orthogonal mRNA orthogonal rRNA pair of  claim 1 , wherein said pair comprises mRNA and rRNA sequences selected from the pairs: 
       
         
           
                 
                 
                 
               
                     
                 
                     
                   Orthogonal mRNA: 
                   Orthogonal mRNA: 
                 
                     
                   Immediately 5′ of AUG 
                   719-726/1531-1542 relative 
                 
                   Pair 
                   initiation codon 
                   to  E.coli  16S rRNA 
                 
                     
                 
                   a 
                   5′-CACCACCCGCAA-3′ 
                   5′-719CCGAGGGC726 (SEQ ID NO: 96) /   
                 
                     
                   (SEQ ID NO: 95) 
                     1531 AUCAAGUGGUUA 1542 -3′ (SEQ ID NO: 97) 
                 
                     
                 
                   h 
                   5′-CACCACCCGCAA-3′ 
                   5′-719CCGCGGGC726 (SEQ ID NO:98)/ 
                 
                     
                   (SEQ ID NO: 95) 
                     1531 AUCACUGUGGUA l542 -3′ (SEQ ID NO: 99) 
                 
                     
                 
                   c 
                   5′-CACCACCCGCAA-3′ 
                   5′-n9CCGCAGGC726 (SEQ ID NO: 100)/ 
                 
                     
                   (SEQ ID NO: 95) 
                     1531 AUCAUUGUGGUA 1542 -3′ (SEQ ID NO: 101) 
                 
                     
                 
                   d 
                   5′-CACCACCCGCAA-3′ 
                   5′-719CCGGUGGC (SEQ ID NO: 102)/ 
                 
                     
                   (SEQ ID NO: 95) 
                     1531 AUCAUUGUGGUA 1542 -3′ (SEQ ID NO: 101) 
                 
                     
                 
                   e 
                   5′-CAACUGCCCGCAA-3′ 
                   5′-719CCGACGGC726 (SEQ ID NO: 104)/ 
                 
                     
                   (SEQ ID NO: 103) 
                     1531 AUCAAUGCAGUA 1542 -3′ (SEQ ID NO: 105) 
                 
                     
                 
                   f 
                   5′-CAACUGCCCGCAA-3′ 
                   5′-n9CCGACGGC726 (SEQ ID NO: 104)/ 
                 
                     
                   (SEQ ID NO: 103) 
                     1531 AUCALJUGCAGUA 1542 -3′ (SEQ ID NO: 106) 
                 
                     
                 
                   g 
                   5′-CAACUGCCCGCAA-3′ 
                   5′-719CCGCGGGC726 (SEQ ID NO: 107/) 
                 
                     
                   (SEQ ID NO: 103) 
                     1531 AUCAUCGCAGUA 1542 -3′ (SEQ ID NO: 108) 
                 
                     
                 
                   h 
                   5′-CAACUGCCCGCAA-3' 
                   5′-719CCGCAGGC726 (SEQ ID NO: 109)/ 
                 
                     
                   (SEQ ID NO: 103) 
                     1531 AUCACCGCAGUA 1542 -3′ (SEQ ID NO: 110) 
                 
                     
                 
                   
                     
                   
                   5′-CAUCCCUCCGCAA-3′ 
                   5′-719CCGCAGGC726 (SEQ ID NO: 109)/ 
                 
                     
                   (SEQ ID NO: 111) 
                     1531 AUCAUGGGAUUA 1542 -3′ (SEQ ID NO: 112) 
                 
                     
                 
                   j 
                   5′-CAUCCCUCCGCAA-3′ 
                   5′-719CCGGUGGC726 (SEQ ID NO: 113)/ 
                 
                     
                   (SEQ ID NO: 111) 
                     1531 AUCAUGGGAUUA 1542 -3′ (SEQ ID NO: 112) 
                 
                     
                 
                       indicates data missing or illegible when filed 
                 
             
                
                
                
                
                
               
               
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         8 . A fusion polypeptide comprising a positive selectable marker polypeptide and a negative selectable marker polypeptide, wherein expression of said fusion polypeptide permits cell survival in the presence of a positive selectable marker and renders cells sensitive to killing by said negative selectable marker. 
     
     
         9 . The fusion polypeptide of  claim 8 , wherein said positive selectable marker comprises an antibiotic resistance coding sequence. 
     
     
         10 . The fusion polypeptide of  claim 9 , wherein said antibiotic resistance coding sequence encodes chloramphenicol acetyltransferase. 
     
     
         11 . The fusion polypeptide of  claim 8 , wherein said negative selectable marker comprises sequence encoding uracil phosphoribosyltransferase. 
     
     
         12 . The fusion polypeptide of  claim 8 , wherein said positive selectable marker comprises chloramphenicol acetyltransferase and said negative selectable marker comprises uracil phosphoribosyltransferase. 
     
     
         13 . The fusion polypeptide of  claim 8 , wherein éxpression of said fusion polypeptide in a cell permits said cell to survive in the presence of chloramphenicol and renders said cell sensitive to killing with 5-fluorouracil. 
     
     
         14 . The fusion polypeptide of  claim 12 , wherein said uracil phosphoribosyltransferase is fused C-terminal to said chloramphenicol acetyltransferase. 
     
     
         15 . A vector comprising a sequence encoding a fusion polypeptide of  claim 8 . 
     
     
         16 . The vector of  claim 15 , wherein said sequence encoding a fusion polypeptide is operably linked to a sequence encoding a ribosome binding sequence. 
     
     
         17 . The vector of  claim 16 , wherein said ribosome binding sequence is a mutated ribosome binding sequence. 
     
     
         18 . The vector of  claim 17 , wherein said ribosome binding sequence comprises a mutation between −13 and +1 relative to an AUG initiation codon at the start of said sequence encoding a fusion polypeptide. 
     
     
         19 . A host cell comprising a vector of  claim 15 . 
     
     
         20 . A cellular logic circuit comprising an orthogonal mRNA orthogonal ribosome pair. 
     
     
         21 . The cellular logic circuit of  claim 20 , wherein said circuit comprises a Boolean AND circuit. 
     
     
         22 . The cellular logic circuit of  claim 20 , wherein said circuit comprises a Boolean OR circuit. 
     
     
         23 . The cellular logic circuit of  claim 20 , wherein said orthogonal mRNA orthogonal rRNA pair comprises a pair selected from those described in  claim 6 . 
     
     
         24 . A cellular logic circuit or cascade comprising two or more orthogonal mRNA orthogonal rRNA pairs recited in  claim 6 . 
     
     
         25 . A method of producing a polypeptide of interest, the method comprising: a) providing an orthogonal mRNA orthogonal rRNA pair, wherein said orthogonal mRNA comprises sequence encoding said polypeptide of interest; and b) introducing nucleic acid encoding said pair to a cell, wherein translation of said orthogonal mRNA by a ribosome comprising said orthogonal rRNA results in production of said polypeptide. 
     
     
         26 . The method of  claim 25 , wherein said orthogonal mRNA orthogonal rRNA pair is a pair as described in  claim 6 .

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