US2008145379A1PendingUtilityA1

Recombinant DNA molecules encoding aminopeptidase enzymes and their use in the preparation of vaccines against helminth infections

Assignee: AGRICULTURAL & FOOD RESPriority: May 8, 1992Filed: Nov 2, 2006Published: Jun 19, 2008
Est. expiryMay 8, 2012(expired)· nominal 20-yr term from priority
C12N 9/48C12N 2799/026C07K 14/4354C07K 2319/02C07K 9/00C07K 9/001C07H 13/04A61P 33/10A61P 33/00A61K 2039/55566A61K 39/0003A61K 39/00
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Claims

Abstract

The present invention provides nucleic acid molecules containing nucleotide sequences encoding helminth aminopeptidase enzymes, and antigenic fragments and functionally-equivalent variants thereof, their use in the preparation of vaccines, for use against helminth parasites, and synthetic polypeptides encoded by them.

Claims

exact text as granted — not AI-modified
1 . A synthetic polypeptide comprising an amino acid sequence encoded by a nucleotide sequence which encodes a helminth aminopeptidase or an antigenic portion thereof, said nucleic acid selected from the group consisting of i) any sequence of SEQ ID NOs:1-15 or 19-21, ii) a sequence fully complementary to said sequence of (i), iii) a portion of said sequence of (i) wherein said portion of said sequence encodes an antigenic peptide of said helminth aminopeptidase, and (iv) a sequence which hybridizes with said sequence of (i) or (ii) under conditions of 2 times SSC, 65 C (where SSC=0.15 M NaCl, 0.015 M sodium citrate, pH 7.2) and which has at least 60% identity with said sequence of (i), other than a synthetic polypeptide corresponding to the protein doublet H110D or a synthetic polypeptide selected from: 
       
         
           
                 
                 
               
                   (SEQ ID NO: 25) 
                     
                 
                 
                 
                 
               
                   (a) 
                   Met Gly Tyr Pro Val Val Lys Val Glu Glu Phe, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 26) 
                     
                 
                 
                 
                 
               
                   (b) 
                   Met Gly Phe Pro Val Leu Thr Val Glu Ser, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 27) 
                     
                 
                 
                 
                 
               
                   (c) 
                   Met Gly/Phe Asn Phe Lys Ile Glu/Val Thr/Glu 
                     
                 
                     
                   Ala Gly, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 28) 
                     
                 
                 
                 
                 
               
                   (d) 
                   Met Lys Pro/Glu Thr/Val Lys Asp/Ala Thr/Lys 
                     
                 
                     
                   Leu - Ile Thr, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 29) 
                     
                 
                 
                 
                 
               
                   (e) 
                   Met Leu Ala Leu Asp Tyr His Ser - Phe Val, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 30) 
                     
                 
                 
                 
                 
               
                   (f) 
                   Met Leu Ala Glu/Tyr Asp Gln/Ala Glu Asp Val, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 31) 
                     
                 
                 
                 
                 
               
                   (g) 
                   Met Gly Phe Pro Leu Val Thr Val Glu Ala Phe 
                     
                 
                     
                   Tyr, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 32) 
                     
                 
                 
                 
                 
               
                   (h) 
                   Met Lys Thr Pro Glu Phe Ala Val/Leu Gln Ala 
                     
                 
                     
                   Phe/Thr Ala Thr Ser/Gly Phe Pro, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 33) 
                     
                 
                 
                 
                 
               
                   (i) 
                   Lys His/Tyr Asn/Val Ser Pro Ala Ala Glu Asn/ 
                     
                 
                     
                   Leu Leu Asn/Gly, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 34) 
                     
                 
                 
                 
                 
               
                   (j) 
                   Lys - Thr Ser Val Ala Glu Ala Phe Asn, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 35) 
                     
                 
                 
                 
                 
               
                   (k) 
                   Lys Ala Ala Glu Val Ala Glu Ala Phe Asp - Ile 
                     
                 
                     
                   - - - Lys Gly, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 36) 
                     
                 
                 
                 
                 
               
                   (l) 
                   Lys Ala Val Glu Val/Pro Ala Glu Ala Phe Asp 
                     
                 
                     
                   Asp Ile Thr? Tyr - - Gly Pro Ser, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 37) 
                     
                 
                 
                 
                 
               
                   (m) 
                   Lys - Glu Glu Thr Glu Ile Phe Asn Met, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 38) 
                     
                 
                 
                 
                 
               
                   (n) 
                   Lys - - - Pro Phe Asn/Asp Ile Glu Ala Leu, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 39) 
                     
                 
                 
                 
                 
               
                   (o) 
                   Asp Gln Ala Phe Ser Thr Asp Ala Lys, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 40) 
                     
                 
                 
                 
                 
               
                   (p) 
                   Met Gly Tyr Pro Val Val Lys Val Glu Glu Phe - 
                     
                 
                     
                   Ala Thr Ala Leu, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 41) 
                     
                 
                 
                 
                 
               
                   (q) 
                   Met Gly Phe Pro Val Leu Thr Val Glu Ser - 
                     
                 
                     
                   Tyr? - Thr, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 42) 
                     
                 
                 
                 
                 
               
                   (r) 
                   Met Glu/Phe Asn Phe Leu Ile Glu/Val Thr/Glu 
                     
                 
                     
                   Ala Gly - Ile Thr, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 43) 
                     
                 
                 
                 
                 
               
                   (s) 
                   Met Gly Phe Leu Val Thr Val Glu Ala Phe Tyr - 
                     
                 
                     
                   Thr Ser, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 44) 
                     
                 
                 
                 
                 
               
                   (t) 
                   Met Lys Thr Pro Glu Phe Ala Val/Leu Gln Ala 
                     
                 
                     
                   Phe/Thr Ala Thr Ser/Gly Phe Pro 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 45) 
                     
                 
                 
                 
                 
               
                   (u) 
                   Met Lys Pro/Glu Thr/Val Leu Asp/Ala Thr/Lys 
                     
                 
                     
                   Leu - Ile Thr - Gly, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 46) 
                     
                 
                 
                 
                 
               
                   (v) 
                   Met Leu Ala Leu Asp Tyr His Ser - Phe Val 
                     
                 
                     
                   Gly?, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 47) 
                     
                 
                 
                 
                 
               
                   (w) 
                   Met Leu Ala Glu/Tyr Asp Gln/Ala Glu Asp Val, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 48) 
                     
                 
                 
                 
                 
               
                   (x) 
                   Lys His/Tyr Asn/Val Ser Pro Ala Ala Glu Asn/ 
                     
                 
                     
                   Leu Leu Asn/Gly, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 49) 
                     
                 
                 
                 
                 
               
                   (y) 
                   Lys - Thr Ser Val Ala Glu Ala Phe Asn, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 50) 
                     
                 
                 
                 
                 
               
                   (z) 
                   Lys Ala Ala Glu Val Ala Glu Ala Phe Asp - 
                     
                 
                     
                   lIe - - - Lys Gly, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 51) 
                     
                 
                 
                 
                 
               
                   (aa) 
                   Lys Ala Val Glu Val/Pro Ala Glu Ala Phe Asp 
                     
                 
                     
                   Asp Ile Thr? Tyr - - Gly Pro Ser, 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 52) 
                     
                 
                 
                 
                 
               
                   (bb) 
                   Lys - Glu Gln Thr Glu Ile Phe Asn Met, 
                     
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 53) 
                     
                 
                 
                 
                 
               
                   (cc) 
                   Lys - - - Pro Phe Asn/Asp Ile Glu Ala Leu 
                     
                 
                   and 
                 
                     
                 
                 
                 
               
                   (SEQ ID NO: 54) 
                     
                 
                 
                 
                 
               
                   (dd) 
                   Asp Gln Ala Phe Ser Thr Asp Ala Lys. 
                     
                 
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
                
               
            
             
                
               
            
             
                
                
                
               
            
             
                
               
            
             
                
               
            
           
         
       
     
     
         2 . A synthetic polypeptide comprising an amino acid sequence encoded by a nucleotide sequence which encodes a polypeptide which raises protective antibodies against helminth parasites, which nucleotide sequence incorporates at least one aminopeptidase-encoding sequence, selected from the group consisting of i) any sequence of SEQ ID NOS: 1-15 or 19-21, ii) a sequence fully complementary to said sequence of (i), iii) a portion of said sequence of (i), and iv) a sequence which hybridizes with said sequence of (i) or (ii) under conditions of 2 times SSC, 65 C (where SSC=0.15 M NaCl, 0.015 M sodium citrate, pH 7.2) and which has at least 60% identity with said sequence of (i), other than a synthetic polypeptide corresponding to the protein doublet H110D or a synthetic polypeptide as defined in (a) to (dd) in  claim 1 . 
     
     
         3 . A synthetic polypeptide comprising an amino acid sequence encoded by a nucleotide sequence which corresponds to or which is complementary to one or more sequences selected from the group of sequences consisting of i) M1(SEQ ID NO:1), ii) B1A (SEQ ID NO:2), iii) B1A-3′ (SEQ ID NO:3), iv) B2 (SEQ ID NO:4), v) M1AUS (SEQ ID NO:5), vi) AusB1 (SEQ ID NO:6), vii) 014-015 (2.5 PCR) (SEQ ID NO:7), viii) 014-872 (3.5 PCR clone 2) (SEQ ID NO:8), ix) A-648 (5′ END of B1) (SEQ ID NO:9), x) A-650 (5′-end of 2.5 PCR) (SEQ ID NO:10), xi) α-649 (5′ end of 3.5 PCR) (SEQ ID NO:11), xii) 014-178 (3′ end of AustB1 clone 2) (SEQ ID NO:12), xiii) 014-178 (3′ end of AustB1 clones 3 and 6) (SEQ ID NO:13) xiv) 014-872 (3.5 PCR clone 10) (SEQ ID NO:14), xv) 014-872 (3.5 PCR clone 19) (SEQ ID NO:15), xvi) H11-1 (SEQ ID NO:19), xvii) H11-2 (SEQ ID NO:20), xviii) H11-3 (SEQ ID NO:21), and xix) a sequence which hybridizes with any of said sequences of (i)-(xviii) under conditions of 2 times SSC, 65 C (where SSC=0.15 M NaCl, 0.015 M sodium citrate, pH 7.2) and which has at least 60% identity with said sequence of (i)-(xviii), other than a synthetic polypeptide corresponding to the protein doublet H110D or a synthetic polypeptide as defined in (a) to (dd) in  claim 1 . 
     
     
         4 . A synthetic polypeptide as claimed in  claim 1 ,  2  or  3  substantially free from other  Haemonchus contortus  components. 
     
     
         5 . A synthetic polypeptide as claimed in  claim 1 ,  2  or  3  in the form of a fusion polypeptide, comprising an additional polypeptide fused to said amino acid sequence as defined in any one of  claims 1  to  3 . 
     
     
         6 . A synthetic polypeptide as claimed in  claim 1 ,  2  or  3  linked to an oligosaccharide having the structure: 
       
         
           
           
               
               
           
         
       
     
     
         7 . A synthetic polypeptide as claimed in  claim 6  wherein said oligosaccharide has the structure: 
       
         
           
           
               
               
           
         
       
     
     
         8 . A synthetic polypeptide as claimed in  claim 1 ,  2  or  3  wherein said synthetic polypeptide is a recombinant polypeptide. 
     
     
         9 . A synthetic polypeptide as claimed in  claim 1 ,  2  or  3  prepared by a method comprising the steps:
 a) culturing a prokaryotic or eukaryotic cell containing a nucleic acid molecule comprising at least one nucleotide sequence which encodes a helminth aminopeptidase or an antigenic portion thereof, said nucleic acid selected from the group consisting of i) any sequence of SEQ ID NOs:1-15 or 19-21, ii) a sequence fully complementary to said sequence of (i), iii) a portion of said sequence of (i) wherein said portion of said sequence encodes an antigenic peptide of said helminth aminopeptidase, and (iv) a sequence which hybridizes with said sequence of (i) or (ii) under conditions of 2 times SSC, 65 C (where SSC=0.15 M NaCl, 0.015 M sodium citrate, pH 7.2) and which has at least 60% identity with said sequence of (i), under conditions whereby said synthetic polypeptide is expressed, and   b) recovering said synthetic polypeptide thus produced.   
     
     
         10 . A vaccine composition for stimulating immune responses against helminth parasites in a human or non-human animal, comprising at least one synthetic polypeptide as defined in  claim 1 ,  2  or  3 , or a virus or host cell having inserted therein a nucleic acid molecule encoding said polypeptide, for stimulation of an immune response to polypeptides encoded by the inserted nucleic acid molecule, together with a pharmaceutically acceptable carrier. 
     
     
         11 . A method of stimulating an immune response against helminth parasites in a human or non-human animal comprising administering to said animal a vaccine composition as defined in  claim 10 .

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