US2003124169A1PendingUtilityA1

Osteogenic devices

Priority: Apr 8, 1988Filed: Sep 19, 2001Published: Jul 3, 2003
Est. expiryApr 8, 2008(expired)· nominal 20-yr term from priority
A61P 43/00A61K 38/00A61F 2310/00365A61P 1/02C07K 14/51A61L 27/3604A61L 2430/02A61L 27/227A61L 27/365A61L 27/56A61C 8/0006A61L 27/3608A61L 27/24A61L 27/34A61K 9/0024A61L 27/3654
52
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Claims

Abstract

Disclosed are 1) osteogenic devices comprising a matrix containing osteogenic protein and methods of inducing endochondral bone growth in mammals using the devices; 2) amino acid sequence data, amino acid composition, solubility properties, structural features, homologies and various other data characterizing osteogenic proteins, 3) methods of producing osteogenic proteins using recombinant DNA technology, and 4) osteogenically and chondrogenically active synthetic protein constructs.

Claims

exact text as granted — not AI-modified
What is claimed is:  
     
         1 . An osteogenic device for implantation in a mammal, said device comprising: 
 a biocompatible, in vivo biodegradable matrix defining pores of a dimension sufficient to permit influx, proliferation and differentiation of migratory progenitor cells from the body of said mammal; and    a protein, produced by expression of recombinant DNA in a host cell, comprising one or more polypeptide chains, each of which has an amino acid sequence sufficiently duplicative of the sequence of COP-5 or COP-7 such that said protein is capable of inducing endochondral bone formation in association with said matrix when implanted in a mammal.    
     
     
         2 . A device for implantation in a mammal, said device comprising: 
 a biocompatible, in vivo biodegradable matrix defining pores of a dimension sufficient to permit influx, proliferation and differentiation of migratory progenitor cells from the body of said mammal; and    a protein, produced by expression of recombinant DNA in a host cell, comprising one or more polypeptide chains, each of which has less than about 200 amino acids, in a sequence sufficiently duplicative of the sequence of COP-5 or COP-7 such that said protein is capable of inducing cartilage formation in association with said matrix when implanted in a mammal.    
     
     
         3 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
               
                     
                 
                           10        20        30        40        50 
                     
                 
                   CXXXXLXVXFXDXGWXXWXXXPXGXXAXYCXGXCXXPXXXXXXXXNHAXX 
                 
                     
                 
                           60        70        80        90       100 
                 
                   QXXVXXXNXXXXPXXCCXPXXXXXXXXLXXXXXXXVXLXXYXXMXVXXCX 
                 
                     
                 
                   CX 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
               
            
           
         
         wherein each X independently represents an amino acid.  
       
     
     
         4 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
               
                     
                 
                           10        20        30        40        50 
                     
                 
                        LXVXFXDXGWXXWXXXPXGXXAXYCXGXCXXPXXXXXXXXNHAXX 
                 
                     
                 
                           60        70        80        90       100 
                 
                   QXXVXXXNXXXXPXXCCXPXXXXXXXXLXXXXXXXVXLXXYXXMXVXXCX 
                 
                     
                 
                   CX 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
               
            
           
         
         wherein each X independently represents an amino acid.  
       
     
     
         5 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
               
                     
                 
                           10        20        30        40        50 
                     
                 
                   CKRHPLYVDFRDVGWNDWIVAPPGYHAFYCHGECPFPLADHLNSTNHAIV 
                 
                    RRRS K S S L  QE VIS E FD Y  E A AY MPESMKAS   VI 
                 
                      KE F E K I  DN     L    N  S   Q  ITK F P    TL 
                 
                       Q     A    S      K 
                 
                     
                 
                           60        70        80        90       100 
                 
                   QTLVNSVNPGKIPKACCVPTELSAISMLYLDENENVVLKNYQDMVVEGCG 
                 
                    SI HAI SEQV EP  A EQMNSLAI FFNDQDK I RK EE T DA H 
                 
                       RF    T   S     K DPV V  Y N S     H RN   RS 
                 
                        N    S                      K       P     E 
                 
                     
                 
                   CR 
                 
                    H 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
         wherein, in each position where more than one amino acid is shown, any one of the amino acids shown may be in that position.  
       
     
     
         6 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
               
                     
                 
                           10        20        30        40        50 
                     
                 
                        LYVDFRDVGWNDWIVAPPGYHAFYCHGECPFPLADHLNSTNHAIV 
                 
                         K S S L  QE VIS E FD Y  E A AY MPESMKAS   VI 
                 
                         F E K I  DN     L    N  S   Q  ITK F P    TL 
                 
                             A    S      K 
                 
                     
                 
                           60        70        80        90       100 
                 
                   QTLVNSVNPGKIPKACCVPTELSAISMLYLDENENVVLKNYQDMVVEGCG 
                 
                    SI HAI SEQV EP  A EQMNSLAI FFNDQDK I RK EE T DA H 
                 
                       RF    T   S     K DPV V  Y N S     H RN   RS 
                 
                        N    S                      K        P    E 
                 
                     
                 
                   CR 
                 
                    H 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
         wherein, in each position where more than one amino acid is shown, any one of the amino acids shown may be in that position.  
       
     
     
         7 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                     
                 
                     
                   Vg1 
                   CKKRHLYVEFK-DVGWQNWVIAPQGYMANYCYGECPYPLTE 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   ILNGSN--H-AILQTLVHSIEPED-IPLPCCVPTKMSP 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISMLFYDNNDNVVLRHYENMAVDECGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         8 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                     
                 
                     
                   DPP 
                   CRRHSLYVDFS-DVGWDDWIVAPLGYDAYYCHGKCPFPLAD 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   HFNSTN--H-AVVQTLVNNNNPGK-VPKACCVPTQLDS 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   VAMLYLNDQSTVVLKNYQEMTVVGCGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         9 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                     
                 
                     
                   OP1 
                        LYVSFR-DLGWQDWIIAPEGYAAYYCEGECAFPLNS 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   YMNATN--H-AIVQTLVHFINPET-VPKPCCAPTQLNA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISVLYFDDSSNVILKKYRNMVVRACGCH 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         10 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                                                          −5 
                     
                 
                     
                     
                                                       HQRQA 
                 
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                 
                     
                   OP1 
                   CKKHELYVSFR-DLGWQDWIIAPEGYAAYYCEGECAFPLNS 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   YMNATN--H-AIVQTLVHFINPET-VPKPCCAPTQLNA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISVLYFDDSSNVILKKYRNMVVRACGCH 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         11 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
               
                     
                 
                     
                   1       10         20        30        40 
                     
                 
                   CBMP-2a 
                   CKRHPLYVDFS-DVGWNDWIVAPPGYHAFYCHGECPFPLAD 
                 
                     
                 
                     
                              50        60         70 
                 
                     
                   HLNSTN--H-AIVQTLVNSVNS-K-IPKACCVPTELSA 
                 
                     
                 
                     
                       80        90       100 
                 
                     
                   ISMLYLDENEKVVLKNYQDMVVEGCGCR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         12 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
               
                     
                 
                     
                   1       10         20        30        40 
                     
                 
                   CBMP-2b 
                   CRRHSLYVDFS-DVGWNDWIVAPPGYQAFYCHGDCPFPLAD 
                 
                     
                 
                     
                              50        60         70 
                 
                     
                   HLNSTN--H-AIVQTLVNSVNS-S-IPKACCVPTELSA 
                 
                     
                 
                     
                       80        90       100 
                 
                     
                   ISMLYLDEYDKVVLKNYQEMVVEGCGCR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         13 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
               
                     
                 
                     
                   1       10         20        30        40 
                     
                 
                   CBMP-3 
                   CARRYLKVDFA-DIGWSEWIISPKSFDAYYCSGACQFPMPK 
                 
                     
                 
                     
                              50        60         70 
                 
                     
                   SLKPSN--H-ATIQSIVRAVGVVPGIPEPCCVPEKMSS 
                 
                     
                 
                     
                       80        90       100 
                 
                     
                   LSILFFDENKNVVLKVYPNMTVESCACR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         14 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                     
                 
                     
                   COP1 
                        LYVDFQRDVGWDDWIIAPVDFDAYYCSGACQFPSAD 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   HFNSTN--H-AVVQTLVNNMNPGK-VPKPCCVPTELSA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISMLYLDENSTVVLKNYQEMTVVGCGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         15 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                     
                 
                     
                   COP3 
                        LYVDFQRDVGWDDWIVAPPGYQAFYCSGACQFPSAD 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   HFNSTN--H-AVVQTLVNNMNPGK-VPKPCCVPTELSA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         16 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                     
                 
                     
                   COP4 
                        LYVDFS-DVGWDDWIVAPPGYQAFYCSGACQFPSAD 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   HFNSTN--H-AVVQTLVNNMNPGK-VPKPCCVPTELSA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         17 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                     
                 
                     
                   COP5 
                        LYVDFS-DVGWDDWIVAPPGYQAFYCHGECPFPLAD 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   HFNSTN--H-AVVQTLVNSVNSKI--PKACCVPTELSA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         18 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                     
                 
                     
                   COP7 
                        LYVDFS-DVGWNDWIVAPPGYHAFYCHGECPFPLAD 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   HLNSTN--H-AVVQTLVNSVNSKI-PKACCVPTELSA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         19 . The device of  claim 1  or  2  wherein the sequence comprises:  
       
         
           
                 
                 
                 
               
                     
                 
                     
                                                  10 
                     
                 
                     
                                               PKHHSQRARKKNKN 
                 
                     
                 
                     
                   1       10         20        30        40 
                 
                   COP16 
                   CRRHSLYVDFS-DVGWNDWIVAPPGYQAFYCHGECPFPLAD 
                 
                     
                 
                     
                              50        60         70 
                 
                     
                   HFNSTN--H-AVVQTLVNSVNSKI--PKACCVPTELSA 
                 
                     
                 
                     
                       80        90       100 
                 
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         20 . The device of  claim 1  or  2  wherein the osteogenics protein comprises a pair of separate polypeptide chains.  
     
     
         21 . Osteogenic protein, produced by expression of recombinant DNA in a host cell, capable of inducing endochondral bone formation in association with a matrix when implanted in a mammal.  
     
     
         22 . A protein, produced by expression of recombinant DNA in a host cell, comprising one or more polypeptide chains less than about 200 amino acids long in a sequence sufficiently duplicative of the sequence of COP-5 or COP-7 such that said protein is capable of inducing cartilage formation in association with a matrix when implanted in a mammal.  
     
     
         23 . The osteogenic protein of  claim 21  having an apparent molecular weight of about 30 kD when oxidized as determined by comparison to molecular weight standards in SDS-polyacrylamide gel.  
     
     
         24 . The osteogenic protein of  claim 23  further characterized by being glycosylated.  
     
     
         25 . The osteogenic protein of  claim 21  having an apparent molecular weight of about 27 kD as determined by comparison to molecular weight standards in SDS-polyacrylamide gel electrophoresis.  
     
     
         26 . The protein of  claim 22  or  25  further characterized by being unglycosylated.  
     
     
         27 . The protein of  claim 21  or  22  comprising a pair of separate polypeptide chains.  
     
     
         28 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
               
                     
                 
                            10        20        30        40 
                     
                 
                   CXXXXLXVXFXDXGWXXWXXXPXGXXAXYCXGXCXXPXXXXXXXXNHAX 
                 
                     
                 
                   50       60        70        80        90 
                 
                   XQXXVXXXNXXXXPXXCCXPXXXXXXXXLXXXXXXXVXLXXYXXMXVX 
                 
                     
                 
                   100 
                 
                   XCXCX 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
         wherein each X independently represents an amino acid.  
       
     
     
         29 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
               
                     
                 
                           10        20        30        40        50 
                     
                 
                        LXVXFXDXGWXXWXXXPXGXXAXYCXGXCXXPXXXXXXXXNHAXX 
                 
                     
                 
                           60        70        80        90       100 
                 
                   QXXVXXXNXXXXPXXCCXPXXXXXXXXLXXXXXXXVXLXXYXXMXVXXCX 
                 
                     
                 
                   CX 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
               
            
           
         
         wherein each X independently represents an amino acid.  
       
     
     
         30 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
               
                     
                 
                           10        20        30        40        50 
                     
                 
                   CKPHPLYVDFRDVGWNDWIVAPPGYHAFYCHGECPFPLADHLNSTNHAIV 
                 
                    RRRS K S S L  QE VIS E FD Y  E A AY MPESMKAS   VI 
                 
                      KE F E K I  DN     L    N  S   Q  ITK F P    TL 
                 
                       Q     A    S      K 
                 
                     
                 
                           60        70        80        90       100 
                 
                   QTLVNSVNPGKIPKACCVPTELSAISMLYLDENENVVLKNYQDMVVEGCG 
                 
                    SI HAI SEQV EP  A EQMNSLAI FFNDQDK I RK EE T DA H 
                 
                       RF    T   S     K DPV V  Y N S     H RN   RS 
                 
                        N    S                      K       P     E 
                 
                     
                 
                   CR 
                 
                    H 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
         wherein, in each position where more than one amino acid is shown, any one of the amino acids shown may be in that position.  
       
     
     
         31 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
               
                     
                 
                           10        20        30        40        50 
                 
                        LYVDFRDVGWNDWIVAPPGYHAFYCHGECPFPLADHLNSTNHAIV 
                 
                         K S S L  QE VIS E FD Y  E A AY MPESMKAS   VI 
                 
                         F E K I  DN     L    N  S   Q  ITK F P    TL 
                 
                             A    S      K 
                 
                     
                 
                           60        70        80        90       100 
                 
                   QTLVNSVNPGKIPKACCVPTELSAISMLYLDENENVVLKNYQDMVVEGCG 
                 
                    SI HAI SEQV EP  A EQMNSLAI FFNDQDK I RK EE T DA H 
                 
                       RF    T   S     K DPV V  Y N S     H RN   RS 
                 
                        N    S                      K       P     E 
                 
                   CR 
                 
                    H 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
         wherein, in each position where more than one amino acid is shown, any one of the amino acids shown may be in that position.  
       
     
     
         32 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                 
                     
                   Vg1 
                   CKKRHLYVEFK-DVGWQNWVIAPQGYMANYCYGECPYPLTE 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   ILNGSN--H-AILQTLVHSIEPED-IPLPCCVPTKMSP 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISMLFYDNNDNVVLRHYENMAVDECGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         33 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                 
                     
                   DPP 
                   CRRHSLYVDFS-DVGWDDWIVAPLGYDAYYCHGKCPFPLAD 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   HFNSTN--H-AVVQTLVNNNNPGK-VPKACCVPTQLDS 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   VAMLYLNDQSTVVLKNYQEMTVVGCGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         34 . The protein of  claim 21  or  22  comprising the amino acid sequence:  
       
         
           
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                 
                     
                   OP1 
                        LYVSFR-DLGWQDWIIAPEGYAAYYCEGECAFPLNS 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   YMNATN--H-AIVQTLVHFINPET-VPKPCCAPTQLNA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISVLYFDDSSNVILKKYRNMVVRACGCH 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         35 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
                 
               
                     
                     
                 
                     
                     
                                                       −5 
                 
                     
                     
                                                       HQRQA 
                 
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                 
                     
                   OP1 
                   CKKHELYVSFR-DLGWQDWIIAPEGYAAYYCEGECAFPLNS 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   YMNATN--H-AIVQTLVHFINPET-VPKPCCAPTQLNA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISVLYFDDSSNVILKKYRNMVVRACGCH 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         36 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
               
                     
                 
                     
                   1       10         20        30        40 
                 
                   CMP-2a 
                   CKRHPLYVDFS-DVGWNDWIVAPPGYHAFYCHGECPFPLAD 
                 
                     
                 
                     
                              50        60         70 
                 
                     
                   HLNSTN--H-AIVQTLVNSVNS-K-IPKACCVPTELSA 
                 
                     
                 
                     
                       80        90       100 
                 
                     
                   ISMLYLDENEKVVLKNYQDMVVEGCGCR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         37 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
               
                     
                 
                     
                   1       10         20        30        40 
                 
                   CBMP-2b 
                   CRRHSLYVDFS-DVGWNDWIVAPPGYQAFYCHGDCPFPLAD 
                 
                     
                 
                     
                              50        60         70 
                 
                     
                   HLNSTN--H-AIVQTLVNSVNS-S-IPKACCVPTELSA 
                 
                     
                 
                     
                       80        90       100 
                 
                     
                   ISMLYLDEYDKVVLKNYQEMVVEGCGCR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         38 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
               
                     
                 
                     
                   1       10         20        30        40 
                 
                   CBMP-3 
                   CARRYLKVDFA-DIGWSEWIISPKSFDAYYCSGACQFPMPK 
                 
                     
                 
                     
                              50        60         70 
                 
                     
                   SLKPSN--H-ATIQSIVRAVGVVPGIPEPCCVPEKMSS 
                 
                     
                 
                     
                       80        90       100 
                 
                     
                   LSILFFDENKNVVLKVYPNMTVESCACR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         39 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                 
                     
                   COP1 
                        LYVDFQRDVGWDDWIIAPVDFDAYYCSGACQFPSAD 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   HFNSTN--H-AVVQTLVNNMNPGK-VPKPCCVPTELSA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISMLYLDENSTVVLKNYQEMTVVGCGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         40 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                 
                     
                   COP3 
                        LYVDFQRDVGWDDWIVAPPGYQAFYCSGACQFPSAD 
                 
                     
                     
                 
                     
                     
                              50        60         70 
                 
                     
                     
                   HFNSTN--H-AVVQTLVNNMNPGK-VPKPCCVPTELSA 
                 
                     
                     
                 
                     
                     
                       80        90       100 
                 
                     
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         41 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
                 
               
                     
                 
                     
                   1        10          20         30         40 
                     
                 
                   COP4 
                         LYVDFS-DVGWDDWIVAPPGYQAFYCSGACQFPSAD 
                 
                     
                 
                     
                               50         60          70 
                 
                     
                   HFNSTN--H-AVVQTLVNNMNPGK-VPKPCCVPTELSA 
                 
                     
                 
                     
                        80         90        100 
                 
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         42 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
                 
               
                     
                 
                     
                   1        10          20         30         40 
                     
                 
                   COP5 
                         LYVDFS-DVGWDDWIVAPPGYQAFYCHGECPFPLAD 
                 
                     
                 
                     
                               50         60          70 
                 
                     
                   HFNSTN--H-AVVQTLVNSVNSKI--PKACCVPTELSA 
                 
                     
                 
                     
                        80         90        100 
                 
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         43 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
                 
               
                     
                 
                     
                   1        10          20         30         40 
                     
                 
                   COP7 
                         LYVDFS-DVGWNDWIVAPPGYHAFYCHGECPFPLAD 
                 
                     
                 
                     
                               50         60         70 
                 
                     
                   HLNSTN--H-AVVQTLVNSVNSKI--PKACCVPTELSA 
                 
                     
                 
                     
                        80         90        100 
                 
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         44 . The protein of  claim 21  or  22  comprising the amino acid sequences:  
       
         
           
                 
                 
                 
               
                     
                 
                     
                                                 −10 
                     
                 
                     
                                              PKHHSSRARKKNKN 
                 
                     
                 
                     
                   1       10        20        30        40 
                 
                   COP16 
                   CRRHSLYVDFS-DVGWNDWIVAPPGYQAFYCHGECPFPLAD 
                 
                     
                 
                     
                               50         60          70 
                 
                     
                   HFNSTN--H-AVVQTLVNSVNSKI--PKACCVPTELSA 
                 
                     
                 
                     
                        80         90        100 
                 
                     
                   ISMLYLDENEKVVLKNYQEMVVEGCGCR 
                 
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         45 . The protein of  claim 21  or  22  comprising the product of expression of a DNA in a procaryotic cell.  
     
     
         46 . A DNA sequence encoding an amino acid sequence sufficiently duplicative of that of the sequence encoded by the gene of Figure  1 A_ such that said encoded sequence induces bone or cartilage formation when implanted in a mammal in association with a matrix.  
     
     
         47 . The DNA of  claim 46  encoding the same amino acid sequence as the gene set forth in FIG. 1A.  
     
     
         48 . The DNA sequence of  claim 46  encoding:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                   1        10         20        30        40 
                     
                 
                     
                   OP1 
                         LYVSFR-DLGWQDWIIAPEGYAAYYCEGECAFPLNS 
                 
                     
                     
                 
                     
                     
                               50         60          70 
                 
                     
                     
                   YMNATN--H-AIVQTLVHFINPET-VPKPCCAPTQLNA 
                 
                     
                     
                 
                     
                     
                        80         90       100 
                 
                     
                     
                   ISVLYFDDSSNVILKKYRNMVVRACGCH 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         49 . The DNA sequence of  claim 46  encoding:  
       
         
           
                 
                 
                 
                 
               
                     
                     
                 
                     
                     
                                                  −5 
                     
                 
                     
                     
                                                   HQRQA 
                 
                     
                     
                 
                     
                     
                   1       10         20        30        40 
                 
                     
                   OP1 
                   CKKHELYVSFR-DLGWQDWIIAPEGYAAYYCEGECAFPLNS 
                 
                     
                     
                 
                     
                     
                               50         60          70 
                 
                     
                     
                   YMNATN--H-AIVQTLVHFINPET-VPKPCCAPTQLNA 
                 
                     
                     
                 
                     
                     
                        80         90        100 
                 
                     
                     
                   ISVLYFDDSSNVILKKYRNMVVRACGCH 
                 
                     
                     
                 
             
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         50 . A cell line engineered to express the protein of  claim 21  or  22 .  
     
     
         51 . The protein of  claim 21  having a half maximum bone forming activity of about 20-25 ng per 25 mg of implant.  
     
     
         52 . A biocompatible, in vivo biodegradable deglycosylated collagenous matrix defining pores of dimensions sufficient to permit influx, proliferation, and differentiation of migratory progenitor cells from the body of a mammal.  
     
     
         53 . The matrix of  claim 52  comprising close-packed particulate matter having a particle size within the range of 70-850 mm.  
     
     
         54 . The matrix of  claim 53  wherein said particulate matter has a particle size within the range of 70-420 mm.  
     
     
         55 . The matrix of  claim 52  defining a shape to span a non-union fracture in said mammal.  
     
     
         56 . The matrix of  claim 52  comprising demineralized, protein-extracted, deglycosylated, particulate xenogenic bone.  
     
     
         57 . The matrix of  claim 52  comprising a material selected from the group consisting of hydroxyapatite, tricalcium phosphate, polymers comprising lactic acid monomer units, polymers comprising glycolic acid monomer units, demineralized, guanidine-extracted, deglycosylated xenogenic bone, and mixtures thereof.  
     
     
         58 . An osteogenic device for implantation in a mammal, said device comprising: 
 a biocompatible, in vivo biodegradable matrix defining pores of a dimension sufficient to permit influx, proliferation and differentiation of migratory progenitor cells from the body of said mammal; and    substantially pure osteogenic protein capable of inducing endochondral bone formation in said mammal disposed in said matrix and accessible to said cells.    
     
     
         59 . The device of  claim 1 ,  2 , or  58  wherein said matrix comprises close-packed particulate matter having a particle size within the range of 70-850 mm.  
     
     
         60 . The device of  claim 1 ,  2 , or  58  wherein said particulate matter has a particle size within the range of 70-420 mm.  
     
     
         61 . The device of  claim 1 ,  2 , or  58  wherein said matrix comprises demineralized, protein-extracted, particulate, allogenic bone.  
     
     
         62 . The device of  claim 1 ,  2 , or  58  wherein said matrix comprises a material selected from the group consisting of collagen, hydroxyapatite, tricalcium phosphate, polymers comprising lactic acid monomer units, polymers comprising glycolic acid monomer units, demineralized, guanidine-eitracted allogenic bone, and mixtures thereof.  
     
     
         63 . The device of  claim 1 ,  2 , or  58  wherein said matrix is shaped to span a non-union fracture in said mammal.  
     
     
         64 . The device of  claim 1 ,  2 , or  58  disposed within the marrow cavity of allogenic bone.  
     
     
         65 . The device of  claim 1 ,  2 , or  58  wherein said matrix comprises demineralized, protein extracted, particulate, deglycosylated zenogeneic bone.  
     
     
         66 . The device of  claim 65  wherein said matrix is treated with a protease.  
     
     
         67 . The device of  claim 58  wherein said osteogenic,protein is unglycosylated.  
     
     
         68 . The device of  claim 67  wherein said osteogenic protein has at apparent molecular weight of about 27 kD when oxidized as determined by comparison to molecular weight standards in SDS-polyacrylamide gel electrophoresis.  
     
     
         69 . The device of  claim 58  wherein said osteogenic protein is glycosylated.  
     
     
         70 . The device of  claim 69  wherein said osteogenic protein has an apparent molecular weight of about 30 kD when oxidized as determined by comparison to molecular weight standards in SDS-polyacrylamide gel electrophoresis.  
     
     
         71 . The device of  claim 58  wherein said osteogenic protein comprises a pair of ppolypeptide chains.  
     
     
         72 . The device of  claim 71  wherein one chain of said pair of polypeptide chains has an apparent molecular weight of about 14 kD and the other has an apparent molecular weight of about 16 kD, both as determined after reduction by comparison to molecular weight standards in SDS-polyacrylamide gel electrophoresis.  
     
     
         73 . The device of  claim 71  wherein one chain of said pair of polypeptide chains has an apparent molecular weight of about 16 kD and the other has an apparent molecular weight of about 18 kD, both as determined after reduction by comparison to molecular weight standards in SDS-polyacrylamide gel electrophoresis.  
     
     
         74 . The device of  claim 58  wherein said osteogenic protein has the approximate amino acid composition set forth below:  
       
         
           
                 
                 
                 
                 
               
                     
                 
                     
                 
                   Amino acid 
                   Rel. no. 
                   Amino acid 
                   Rel. no. 
                 
                   residue 
                   res./molec. 
                   residue 
                   res./molec. 
                 
                     
                 
                   Aspartic acid/ 
                   22 
                   Tyrosine 
                   11 
                 
                   Asparagine 
                     
                   Valine 
                   14 
                 
                   Glutamic acid/ 
                   24 
                   Methionine 
                    3 
                 
                   Glutamine 
                     
                   Cysteine 
                   16 
                 
                   Serine 
                   24 
                   Isoleucine 
                   15 
                 
                   Glycine 
                   29 
                   Leucine 
                   15 
                 
                   Histidine 
                    5 
                   Proline 
                   14 
                 
                   Arginine 
                   13 
                   Phenylalanine 
                    7 
                 
                   Threonine 
                   11 
                   Tryptophan 
                   ND 
                 
                   Alanine 
                   18 
                 
                   Lysine 
                   12 
                 
                     
                 
                     
                 
             
                
                
                
                
                
               
               
                
                
                
                
                
                
                
                
                
                
                
                
                
               
            
           
         
       
     
     
         75 . The device of  claim 58  wherein said osteogenic protein comprises the amino acid sequence: 
 VPKPCCAPT  
 
     
     
         76 . The device of  claim 1  or  58  wherein the half maximum bone inducing activity of said protein is 0.8 to 1.0 ng per mg of said matrix.  
     
     
         77 . A method of inducing local cartilage or bone formation in a mammal comprising the step of implanting the device of  claim 1 ,  2 , or  58  in said mammal at a locus accessible to migratory progenitor cells of said mammal.  
     
     
         78 . A method of inducing endochondral bone formation in a mammal comprising the step of implanting the device of  claim 1  or  58  in said mammal at a locus accessible to migratory progenitor cells of said mammal.  
     
     
         79 . A method of inducing endochondral bone formation in a non-union fracture in a mammal comprising the step of implanting in the fracture in said mammal the device of  claim 63 .  
     
     
         80 . Antibodies reactive with an epitope of the protein of  claim 21  or  22 .

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